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Zoology Publications from Victoria University of Wellington—Nos. 68, 69 and 70

Keys to Aid in the Identification of Marine Harpacticoid Copepods. Amendment Bulletin No. 1

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page 1

Keys to Aid in the Identification of Marine
Harpacticoid Copepods
 Amendment Bulletin No. 1.

by
J. B. J. Wells,
Victoria University of Wellington.

Introduction

The purpose of this bulletin is to illustrate one way in which information that has appeared in the literature since publication of my Keys (Wells, 1976) can be incorporated in them. Also, since the Keys were designed to be useful to non-specialists in harpacticoid taxonomy such bulletins (I hope to publish them at regular intervals) may serve to bring to the user's attention all the recent taxonomic literature. In this first bulletin I take the opportunity of correcting errors in the Keys. Some I have noticed myself but my attention has been drawn to others by several colleagues; to them I extend my grateful thanks. Hopefully few now remain but I would welcome notice of any that have as yet gone undetected.

In this bulletin the word codon is introduced as the term for the set of character states that characterize a taxon or group of taxa.

Family Canuellidae

1.There are two errors in the Key, correct as follows (p.19):
Brianola elegans 6:4:4:4/6:5:4:4/a/3/6
B. sydneyensis 5:4:4:4/6:5:4:4/a/3/6
2.General Notes (p.20): The most recent revision of this family is by Por, 1967, not Por, 1969.

Family Ectinosomidae

1.Family name must be changed to Ectinosomatidae (pp.27-36); the faulty latinization is pointed out by Moore (1976).
2.KGG 1: Arenosetella monensis Moore, 1976, to genus codon (p.28).
3.KGG 100: Ectinosoma soyeri Apostolov, 1975, to genus codon (p.29).
4.KGG 300 (p.33):
(a) Hastigerella soyeri Bodin, 1976, nom. nov. for H. noodti Soyer, 1974.
(b) Hastigerella bodini Apostolov, 1974, to codon for Hastigerella a).
5.KGG 500: Noodtiella wellsi Apostolov, 1974, to genus codon (p.35).
6.General Notes (p.36): Pallares (1975b) redescribes the female of Halophytophilus fusiformis.

Family D'Arcythompsoniidae

As Articles 27 and 32(c) of the International Code of Zoological Nomenclature forbid the use of an apostrophe D'Arcythompsonia T. Scott must be corrected to Darcythompsonia and the family name changed accordingly. Corrections are required on pp.6, 8, 37, 38 and 39.

page 2

Family Harpacticidae

1.KGG 1 (p.43): Perissocope exiguus Pallares, 1975d, new codon— 2/3:3:3/p/7:8:7/5:6:5.
2.KGG 100 (p.45):
(a) Harpacticus nipponicus Ito, 1976, to codon for Harpacticus c).
(b) Harpacticus septentrionalis yamadai Ito, 1976; add superscripta) to the species in footnote b) on p.46 and add the subspecies to item 2 in General Notes (p.47).
(c) Harpacticella paradoxa is redescribed by Ito & Kikuchi (1977); delete from footnote d) (p.46) and add new codon—nor /7/1♀/5:5/0-1:5.

Family Tisbidae

1. Tisbe varipes Marcus, 1974a, to genus codon in KGG 100 (p.49).
2.For the genus Scutellidium (KGG 100, p.49 and General Notes, p.52) note this information:
(a)Branch (1975) suggests—
S. spinatum is a synonym of S. ringueleti,
S. purpurocinctum is a synonym of S. arthuri,
S. loureiroi and S. deseadensis possibly are synonymous with S. lamellipes, which itself may be a synonym of S. longicauda.
(b) S. arthuri is redescribed by Ito, 1976.
(c) S. macrosetum Branch, 1975, S. caeneus Ito, 1976, S. hirutai Ito, 1976, and S. longicauda acheloides Ito, 1976, all to genus codon in KGG 100.
(d) S. boreale Ito, 1976 and S. hippolytes akaba Ito, 1976 to codon for S. hippolytes in KGG 100; amend footnote c) appropriately.

Family Peltidiidae

1.KGG 1 (p.53):
(a) Neopeltopsis pectinipes Hicks, 1976, new codon—
1/2/0:0:0/1:1:1/d:d.
(b) Eupelte acutispinis Zhang & Li, 1976, new codon—
2/2/0:0:0/2:2:1/d:d.
2.KGG 100: Paralteutha minuta Ramirez, 1971, to genus codon (p.55).
3.p.55: Note that Paralteutha villosa is redescribed by Pallares (1975b).

Family Thalestridae

1.KGG 1:
(a) Idomene cookensi Pallares, 1975b, to codon for I. scotti (p.58).
(b) Dactylopodia pectenis Pallares, 1975d to codon for D. pontica (p.59).
(c) Diarthrodes ponticus orientalis Apostolov, 1975; add superscripta) to species in footnote b) (p.60) and add the subspecies to item 2 in General Notes (p.69).
(d) Diarthrodes tetrastachyus Yeatman, 1976, new codon—
2:3/7:8:8/2:2/3/p:f.
page 3
2.KGG 100 (p.61):
(a) Dactylopodia decostata Pallares, 1975d, to genus codon.
(b) Paradactylopodia hexiarticulata Kunz, 1975, to genus codon.
3.KGG 400: Parathalestris patagonica, P. vinosa, P. aurantiaca, all new species by Pallares, 1975d, to genus codon (p.64).
4.General Notes (p.69):
(a)Pallares (1975b) redescribes Parathalestris ganio.
(b)Pallares (1975c) describes the male of Paramenophia platysoma (KGG 300, p.63).

Family Parastenhelidae

The description of Parastenhelia reducta Apostolov, 1975, requires a note of this new species in item 7 on p.12. Also, in the Key to Families, item 47 sub. 5 (p.10) now must read "with 4:4-5:4-5 .......", but note that this species will also key out in item 47 sub. 2 and then in item 49 sub. 2, to Thalestridae.

Family Diosaccidae

1.KGG 100: Robertgurneya soyeri Apostolov, 1974 and Robertgurneya sp. Apostolov, 1973, to codon for R. oligochaeta (p.73).
2.KGG 110: Amphiascoides bulbiseta Pallares, 1975e, to codon for Amphiascoides d) (p.74).
3.KGG 210 (p.78):
(a) Typhlamphiascus typhlops: It is now known that P.4 Enp. 2 has an inner seta in populations from the Maldive Islands, Germany, Sweden and Spitzbergen (Becker, 1970). It is not known if Sars' original description is correct. Consequently the species must be added to footnote a).
(b) Typhlamphiascus lamellifer capensis Kunz, 1975, new codon—
5/5/5:6/?
(also, this species must be added to KGG 200 (p.76) where it keys out with T. typhlops).
4.KGG 230 and 240: Amphiascus elongatus Ito, 1972: The present codon in KGG 230 (p.80) applies to the female only. The species requires a new codon in KGG 240 (p.81) —
tr/sw/5-6:6/2:6.
5.KGG 240: Amphiascus longarticulatus Marcus, 1974b, to codon for A. tenuiremis and A. gracilis (p.81).
6.KGG 260: Paramphiascopis triarticulatus Moore, 1976, to genus codon (p.84).
7.KGG 400 (p.86):
(a) Stenhelia (D.) oblonga and S. (D.) elisabethae: Codons for these species are wrong. The correct codon for both is—
4/7:8:8/8/d:f/lss.
(b) S. (D.) normani: Codon on p.86 applies only to subspecies normani. The codon for the other subspecies (n. polluta and n. acutirostris) is—
4/7:8:8/8/d:f/lss.
(c) S. (D.) bifida Coull, 1976, new codon—
5/6:6:7/8/d:d/lss.
page 4
8.KGG 500 (p.88): Stenhelia (D.) palustris bispinosa Bodin, 1970, new codon—
3:3:2/6:7:6/0:0:0.
Also, indicate in KGG 400 (p.86) that the codon refers to the nominate subspecies only.

Family Ameiridae

1. Leptomesochra sp. Bodin, 1970 is identified by Bodin (1972) as Mesochra pygmaea; delete from KGG 1 (p.102).
2.KGG 1:
(a) Karllangia tertia Kunz, 1975, to codon for K. psammophila (p.101).
(b) Parapseudoleptomesochra dubia Kunz, 1975, new codon (p. 102) —
3:3/3:3:3/3:3:2/4/0.
3.KGG 300 (pp.107-108):
(a) Sarsameira elegantula Kunz, 1975, new codon—
s/1:1:1/1:1:1/4:4:4/2.
(b) Stenocopia longiseta: Kunz (1975) describes the female (see footnote a).
(c) Nitocrella stygia Apostolov, 1976 and N. hibernica bulgarica Apostolov, 1976, are not this genus but Nitocra; to codon for Nitocra e).
(d) Psyllocamptus (Langpsyllocamptus) longisetosus Kunz, 1975, new codon—
s/0:0:0/0:0:0/3:3:3/1-2.
4.KGG 330 (p.111): Nitocra sewelli huntsmani Kunz, 1976, new codon—
7:7:7/1:1:1/5:6/3:6,
and indicate in KGG 500 (p.117) that the codon refers to the nominate subspecies only.
5.KGG 400 (p.114):
(a) Ameira usitata is redescribed by Kunz (1975): delete from footnote b) (p. 116) and add new codon—
0:0:0/1:1:1/1:1:1/4:5:5/4:5/3(?):5.
(b) Psyllocamptus (Langpsyllocamptus) quadrispinosus Kunz, 1975, new codon—
0:0:0/0:0:0/0:0:0/3:3:3/5:4/?.
6. Psyllocamptus is divided by Kunz (1975) into two sub-genera:
Psyllocamptus s.str. for monachus (KGG 1) (p.101), carolinensis (KGG 1) (p.101), propinquus (KGG 100) (p.105), minutus (KGG 200) (p.106) and bermudae (KGG 200) (p.106);
Langpsyllocamptus nov. for quinquespinosus (KGG 300) (p.108), longisetosus (KGG 300) (see note 3 above), triarticulatus (KGG 400) (p.114) and quadrispinosus (KGG 400) (see note 5 above). page 5
7. Ameiropsis australis Kunz, 1975: Kunz states that P.1 Exp.2 has a very reduced, hair-like inner seta. As this is not a very obvious structure it may be best to place the species in both KGG 300 and 400, with an appropriate note. In KGG 400 (p.114) place a new codon—
1:1:1/1:1:1/1:1:1/4:5:5/5:5/3:5.
In KGG 300 (p.107) A. australis keys out with Sarsameira exilis and S. peresi, s/1:1:1/1:1:1/4:5:5/2. This codon should now lead to KGG 350, a new KGG defined as follows—
KGG 350 — characters
1.A.1♀
n — number of segments.
2.P.2-P.4 Exp.3
n:n:n — number of setae and/or spines on P.2, P.3 and P.4 respectively.
3.P.5♀
n :n — number of seteae on Benp. and Exp. respectively.
4.P.5♂
n :n — number of setae on Benp. and Exp. respectively.
KGG 350
A.1.♀ segs. P.2-P.4 Exp.3 setae P.5 ♀ setae P.5 ♂ setae
9 7:7:7 5:6 5:6 Sarsameira exilis
9 6:7:8 5:6 ? S. peresi
8 6:6:7 5:5 3:5 Ameiropsis australis Kunz, 1975
8.Petkovski (1976) has revised Nitocrella. He removes some species to Parapseudoleptomesochra and erects two new genera, Nitocrellopsis and Stygonitocrella. He describes three new species of Nitocrella s.str. and follows Borutzky (1969) in placing Sterba's (1968) species Spelaeocamptus asiaticus and S. reductus in Nitocrella s.str. However, it must be noted that Borutzky's renaming of S. reductus as N. sterbai is not valid. Borutzky apparently was unaware that Noodt (1957) has already returned Nitocrella reducta (Schäfer) to the genus Nitocra. Sterba's original name must be restored and N. sterbai lapse as a synonym of N. reducta (Sterba). Petkovski's revision requires the following amendments to the Keys—
(a)KGG 1:
(i) Nitocrella d) (p.102) becomes Parapseudoleptomesochra d) with consequent change to footnote d) (p.104).
(ii) Nitocrella g) and h) (p.103) become Stygonitocrella g) and h) respectively, with consequent changes to the appropriate footnotes (p.104).
(iii)On p.102 the species Nitocrella tridens, botosaneanui, subterranea and heruridensis are removed to Parapseudoleptomesochra and the species Nitocrella intermedia, elegans and ionelli are removed to Nitocrellopsis.page 6
(iv)On p.103 the species Nitocrella karamam, colchica, dubia, insularis, ljovuschkini and orghidani are removed to Stygonitocrella.
(b)KGG 600 (pp. 118-119): The following new codons are to be added—
4:4:6/1:1:1/2:2:2/2-3:4/2 :4 Nitocrella caraioni Petkovski, 1976
4:4:6/1:1:1/2:2:2/0:4/0:5 N. motasiPetkovski, 1976
4:4:5/1:1:1/2:2:2/3:4/2:5 N. cubanorum Petkovski, 1976
4:4:5/1:1:1/2:2:2/0:2/? N. asiatica (Sterba, 1968)
4:4:5/1:1:1/2:2:2/4b)/? N. reducta (Sterba, 1968)
with the footnote—
b) p.5 reduced to a single plate.
(c)General Notes item 6 (p.123): This comment now applies to the four genera Nitocrella, Nitocrellopsis, Stygonitocrella and Parapseudoleptomesochra.

Family Paramesochridae

KGG 1 (p.124):

(a) Caligopsyllus primus Kunz, 1975, new codon—
2:0:0:0/1/na/5/nf.
(b) Kliopsyllus sp. Apostolov, 1973, new codon—
2:1:1:1/3/1/4/?.
(c)The species K. minutus, K. pseudogracilis and K. runtzi (p.125) are all species of Kliopsyllus.

Family Tetragonicipitidae

KGG 1 (p.131):

(a) Phyllopodopsyllus langi Kunz, 1975, new codon—
a/2wd/2/3:2:3/5:4:5.
(b) Phyllopodopsyllus paraborutzkyi Kunz, 1975, new codons—
a/2wd/2/2:3:3/4:4:6 ♀
a/2wd/2/2:3:2/4:4:6 ♂

Family Canthocamptidae

1.KGG 1 (p.135) : Itunella intermedia Apostolov, 1975, new codon—
a/1:4/2:1:1:1/1:1:1/7.
The existing codon for Itunella characterizes the other three species of the genus—tenuiremis, muelleri and bacescui.
2.KGG 100 (p.137): Mesochra bodini Kunz, 1975, new codon—
6/3/5-6:7:7/5:5/5:5/2:6.
page 7

Family Cylindropsyllidae

1. Arenopontia stygia: Lindgren (1976) considers that this species may be only an intra-population variant of A. arenarida (KGG 600) (p.149).
2.KGG 600: Arenopontia riedli Lindgren, 1976, to codon for A. arenarida (p. 149).

Family Cletodidae

1.Error in KGG 500 (p.164): Character 5 should read "P.3 ♂ Enp.".
2.Soyer (1975a) describes three new species of Mesocletodes, all of which can be included in KGG 1 (p.154) —
M. carpinei, new codon—
3:2/3:3:3/2:2/7:7:7/4:4.
M. bodini, new codon—
3:2/3:3:3/2:2/7:7:7/5:5.
M. ameliae, to codon for M. commixtus (p.157).
3. Cletocamptus alburquerquensis: This species is included in Cleto-camptus e) of KGG 1 (p.157) by mistake. Delete from footnote e) (p.159) and add new codon—
3:2/3:3:3/2:1/5:5:4/3:2.
4.KGG 1 (p.154):
(a) Australonannopus aestuarinus Hamond, 1974, new codons—
3:2/3:3:3/2:0/4:4:5/2:na ♀
3:2/3:3:3/2:0/4:4:4/2:na ♂
(b) Paranannopus wellsi Soyer, 1975b, new codon—
3:2/3:3:3/3:2/7:8:8/5:6.
5.KGG 500 (p.165): Pallares (1975e) notes some variation in Enhydrosoma propinquum; add a second codon for this species—
2:2/a/d:3:5/d:2:2/♀.

Family Laophontidae

1. Laophonte sinifer Kunz, 1975, to codon for Paralaophonte spinicauda in KGG 1200-♀ ♀ (p.182) and to codon for Paralaophonte a) in KGG 1100-♂ ♂ (p.197).
2. Pseudocleta corbula: Present codon in error; correct in KGG 1- ♀ ♀ (p. 172) as follows—
3:3:3/1:1:1/5:6/3/6.
3. Heterolaophonte cambelliensis: P.5 ♂ can have a total of 5 or 7 setae. Therefore this species must be added to footnote k) of KGG 1-♂ ♂ (p.191).
4. Microlaophonte trisetosa Boxshall, 1976, new codons—
KGG 1-♀ ♀ (p.172) —
3:3:3/2:2:2/6:6/1/6.
KGG 1-♂ ♂ (p.188) —
3:3:3/2:2/0:4/1/3.
5. Phycolaophonte insularis Pallares, 1975a, new codons—
KGG 1400-♀ ♀ (p.183) —
5:5:5-6/0:0:0/0:0:0/2:3(4):3(4)b)/0:0:0.
(with new footnote — "b) P.3 and P.4 Enp. 2 have 3 setae plus a fused spine").
KGG 1-♂ ♂ (p.190) —
3:3:3/2:2/2:5/2/2. page 8
6. Cletopsyllus bacescui Marcus, 1976:
KGG 1600-♀ ♀ (p.184) — to codon for Pseudocletopsyllus spiniger.
KGG 1-♂ ♂ (p.188), new codon—
3:3:3/2:2/3:4/3/2.
7. Laophonte dinocerata: The male is described by Pallares (1975b); to codon for L. inornata and L. sima in KGG 1300-♂ ♂ (p.198).

Family Ancorabolidae

1.p.203 item 1: Delete entirely; A. s. spinifera was described by Norman in 1911 and appears in Lang's Monograph.
2. Laophontodes whitsoni (p.202) is redescribed by Pallares (1975b) who describes P.4 Enp.2 with only 2 setae. Also, there is an error in the existing key which puts L. whitsoni in a codon with L. bicornis and hamatus. Remove from this codon and replace with this new codon—
3:2/2:2/4/6:7:7/2:2 or 4.

Family Latiremidae

Bodin (1975) has formally proposed that Latiremus Bozic is a synonym of Delamarella Chappuis (see p.12 of the Keys). A simple key to the sole genus of the family can be constructed.

The relationships of the family are not at all clear, although it would appear to belong to the Suprafamily Ameiridimorpha Lang. Further research is required and until then it is best to treat the family as incertae sedis in Harpacticoida and thus to place it at the end of the sequence of families.

KGG 1 — characters

1.A.1♀
n — number of segments.
2.P.1.
n:n — number of segments in Exp. and Enp. respectively.
3.P.1 Exp., distal segment
n — number of setae and/or spines.
4.P.2-P.3 Exp.2
n:n — number of setae on inner border of P.2 and P.3 respectively.
5.P.2-P.3 Enp.3
n:n — number of setae on P.2 and P.3 respectively.
KGG 1
A.1♀ P.1 segs. P.1 Exp. distal segs. setae P.2-P.3 Exp.2 inner setae P.2-P.3 Enp.3 setae
8 2:2 5 1:1 3:3 Delamarella arenicola
9 2:2 5 1:1 2:2 D. karamani
9 2:2 5 0:0 2:2 D. galateae
8 3:2 4 1:1 3:2 D. eximia
page 9

Typographic Errors

A number of typographic errors have been brought to my notice. Corrections should be made as follows:
  • p.18 Scottolana bulbosa (Por, 1964 )
  • p.18 Scottolana bulbifera
  • p.40 Danielssenia stefanssoni (i.e. not stefannsoni)
  • p.44 Perissocope litoralis (i.e. not littoralis)
  • p.52 Tisbe bulbisetosa
  • p.58 Dactylopusioides macrolabris
  • p.60 Diarthrodes campbelliensis
  • p.69 Dactylopodia longyearbyenensis
  • p.76 Robertgurneya (?) arabica
  • p.80 Amphiascus gauthieri
  • p.88 Stenhelia longifurca (i.e. not longifurcata)
  • p.94 Schizopera validior
  • p.99 Robertsonia tenuis kieliensis
  • p.101 Parapseudoleptomesochra reducta
  • p.104 Interleptomesochra eulitoralis (i.e. not littoralis)
  • p.127 Kliopsyllus spiniger Wells, Kunz & Rao, 1975 (bibliography requires consequent amendment)
  • p.128 Kliopsyllus longifurcatus
  • p.129 Apodopsyllus schulzi (i.e. not schultzi)
  • p.132 Phyllopodopsyllus bahamensis (i.e. not bahamaensis)
  • p.141 Sewellina reducta
  • p.145 Leptastacus waltairensis
  • p.146 Paraleptastacus supralitoralis (i.e. not supralittoralis)
  • p.149 Arenopontia nesaie
  • p.150 Arenopontia spinicaudata
  • p.151 Ichnusella pasquinii
  • p.151 Psammopsyllus limnicola
  • p.155 Metahuntemannia spinosa
  • p.156 Eurycletodes verisimilis
  • p.158 Mesopsyllus atargatis (i.e. not atagartis)
  • p.165 Enhydrosoma curticauda (i.e. not curticaudata)
  • p.165 Enhydrosoma buchholtzi
  • p.167 Cletodes carthaginiensis
  • p.180 Laophonte recticaudata.
  • p.181 & 182 Esola spelaea
  • p.186 Heterolaophonte campbelliensis
  • p.196 Laophonte longicaudata
page 10

References

Note that Bodin (1975, 1976) has published two further additional lists to supplement his catalogue of new species.

Apostolov, A. , 1973. Notes sur les harpacticoides (Crustacea, Copepoda) de la Mer Noire. Zoologischer Anzeiger 190: 174-189.

1974 Copépodes harpacticoides de la Mer Noire. Travaux du Muséum d'Histoire Naturelle 'Grigore Antipa' 15: 131-139.

1975. Les harpacticoides marins de la Mer Noire. Description de quelques formes nouvelles. Vie et Milieu 25A: 165-178.

1976. New species of Harpacticoida (Copepoda) from Bulgaria. Zoologichesky Zhurnal 55: 453-458 (Russian text, English summary).

Becker, K. H. , 1970. Beitrag zur Kenntnis der Copepoda Harpacticoida sublitoraler Weichböden in der Kieler Bucht. Kieler Meeresforschungen 26: 56-73.

Bodin, P. , 1970. Copépodes harpacticoides marins des environs de La Rochelle. 1. — Espèces de la vase intertidale de Chatelaillon. Tethys 2: 385-436.

1972. Copépodes harpacticoides marins des environs de La Rochelle. 2. — Espèces de la zone intertidale d'Yves. Tethys 3: 411-433.

1975. Catalogue des nouveaux copépodes harpacticoides marins. Additif No. 2. Tethys 7: 265-278.

1976. Catalogue des nouveaux copépodes harpacticoides marins. Additif No. 3. Université de Bretagne Occidentale, Laboratoire d'Oceanographie Biologique pp.45.

Borutzky, E. V. , 1969. Copepoda Harpacticoida fauna from the subterranean waters of middle Asia in zoogeography aspect. Nauchyne Doklady Vysshei Shkholy, Biologischeskie Nauki (Moscow) 1969 (1): 7-11 (Russian text; title is quoted as it appears in transliteration in Zoological Record 106B (10): 11, 1969. Journal title is the transliteration given in the 1975 Biosis List of Serials, which states that it can be cited interchangeably with Biologischeskie Nauki).

Boxshall, G. A. , 1976. Microlaophonte trisetosa sp. nov., a new harpacticoid copepod from southern Florida. Bulletin of the British Museum (Natural History), (Zoology) 30: 223-228.

Branch, G. M. , 1975. A new species and records of Scutellidium (Copepoda, Harpacticoidea) from South Africa, and a world key to the genus. Annals of the South African Museum 66: 221-232.

Coull, B. C. , 1976. A revised key to Stenhelia (Delavalia) (Copepoda: Harpacticoida) including a new species from South Carolina, U.S.A. Zoological Journal of the Linnean Society of London 59: 353-364.

Hamond, R. , 1974. Australonannopus aestuarinus gen. nov., sp. nov. (Crustacea: Harpacticoida), an aberrant cletodid from Australian brackish waters Papers and Proceedings of the Royal Society of Tasmania 107: 239-245.

Hicks, G. R. F. , 1976. Neopeltopsis pectinipes, a new genus and species of seaweed-dwelling copepod (Harpacticoida: Peltidiidae) from Wellington, New Zealand. New Zealand Journal of Marine and Freshwater Research 10: 363-370.

Ito, T. , 1972. Descriptions and records of marine harpacticoid copepods from Hokkaido, IV. Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology 18: 305-336.

1976: Descriptions and records of marine harpacticoid copepods from Hokkaido, VI. Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology 20: 448-567.

Ito, T. & Kikuchi, Y , 1977. On the occurrence of Harpacticella paradoxa (Brehm) in Japan; a fresh-water harpacticoid copepod originally described from a Chinese lake. Annotationes Zoologicae Japonenses 50: 40-56.

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Kunz, H. , 1975. Copepoda Harpacticoidea aus dem Litoral des südlichen Afrika. 1. Teil. Kieler Meeresforschungen 31: 179-212.

1976. Eine neue Unterart der Gattung Nitocra (Copepoda, Harpacticoidea) aus Norddeutschland. Gewässer und Abwässer 60/61: 27-34.

Lindgren, E. W. , 1976. Five species of Arenopontia (Copepoda, Harpacticoida) from a North Carolina beach, U.S.A. Crustaceana 30: 229-240.

Marcus, A. , 1974a. Tisbe varipes n.sp. (Copepoda, Harpacticoida) de la Mer Noire. Travaux du Muséum d'Histoire Naturelle 'Grigore Antipa' 14: 115-123.

1974b. Contributions to the study of the genus Amphiascus Sars (part.), Copepoda, Harpacticoida from the Black Sea. Travaux du Muséum d'Histoire Naturelle 'Grigore Antipa' 15: 111-122.

1976. Contributions to the study of the genus Cletopsyllus Willey, 1935 (Copepoda, Harpacticoida) from the Indian Ocean. Travaux du Muséum d'Histoire Naturelle 'Grigore Antipa' 17: 39-49.

Moore, C. G. , 1976. The harpacticoid families Ectinosomatidae and Diosaccidae (Crustacea, Copepoda) from the Isle of Man. Journal of Natural History 10: 131-155.

Noodt, W. , 1957. Zur Kenntnis von Nitocra reducta Schafer (Copepoda, Harpacticoidea). Zoologischer Anzeiger 159: 179-184.

Pallares, R. E. , 1975a. Sobre un nuevo genero de la familia Laophontidae. Contrubucion Cientifica del Centro de Investigacion de Biologia Marina, Buenos Aires 93: 1-5.

1975b. Copepodos harpacticoides marins de Tierra del Fuego (Argentina), 1. Isla de los Estados. Contribucion Cientifica del Centro de Investigacion de Biologia Marina, Buenos Aires 122: 1-34.

1975c, Sobre el hallazo de ejemplares masculinos de Paramenophia platysoma (Thompson and A. Scott, 1903) (Copepoda, Harpacticoida). Contribucion Cientifica del Centro de Investigacion de Biologia Marina, Buenos Aires 135: 1-6.

1975d. Copepodos marinos de la Ria Deseado (Santa Cruz, Argentina). Contribucion sistematico-ecologica IV. Physis, Buenos Aires 34A: 67-83.

1975e. Copepodos marinos de la Ria Deseado (Santa Cruz, Argentina). Contribution sistematico-ecologica IV (conclusion). Physis, Buenos Aires 34A: 213-227.

Petkovski, T. K. , 1976. Drei neue Nitocrella-Arten von Kuba, zugleich eine revision des genus Nitocrella Chappuis (s.restr.) (Crustacea, Copepoda, Ameiridae). Acta Musei Macedonici Scientiarium Naturalium 15: 1-26.

Por, F. D. , 1967. Level bottom Harpacticoida (Crustacea, Copepoda) from Elat (Red Sea), part 1. Israel Journal of Zoology 16: 101-165.

Ramirez, F. C. , 1971. 'Paralteutha minuta', una nueva especie de copepodos (Harpacticoida, Peltidiidae) hallado en aguas conteras de Mar del Plata, Argentina. Revista del Museo de La Plata, Zoologia 11: 115-119.

Soyer, J. , 1975a. Contribution a l'étude des copépodes harpacticoides de Méditerranée occidentale. 13. Le genre Mesocletodes Sars (Cletodidae, T. Scott). Systématique, écologie. Vie et Milieu 25B: 157-173.

1975b. Contribution a l'étude des copépodes harpacticoides de Méditerranée occidentale. 14. Le genre Paranannopus Lang (Cletodidae, T. Scott). Systématique, écologie. Vie et Milieu 25B: 331-338.

Sterba, C. , 1968. Neue Harpacticoidea (Crustacea, Copepoda) aus dem asiatischen Teil der Palarktis. Zoologischer Anzeiger 180: 50-68.

Wells, J. B. J. , 1976. Keys to Aid in the Identification of Marine Harpacticoid Copepods. Department of Zoology, University of Aberdeen pp. 215.

Yeatman, H. C. , 1976. Marine littoral copepods from Jamaica. Crustaceana 30: 201-219.

Zhang, C. & Li, Z. , 1976. Harpacticoida (Copepoda, Crustacea) from Xisha Islands of Guangdong Province, China. Acta Zoologica Sinica 22: 66-70. (Chinese text, English summary).

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