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Zoology Publications from Victoria University of Wellington—Nos. 76 and 77

Keys to Aid in the Identification of Marine Harpaticoid Copepods. Amendment Bulletin No. 4

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Keys to Aid in the Identification of Marine Harpacticoid Copepods
Amendment Bulletin No. 4

by
J. B. J. Wells,
Victoria University of Wellington.
Zoology Publications from Victoria University of Wellington No. 77, issued August 1983.

Introduction

This Bulletin deals with several important papers, but particularly notable are the revision of the Paramesochridae by Kunz (1981), the discussion of Oniscopsis by Becker & Kunz (1981) and the partial revision of Tisbe by Volkmann (1979c); all are major contributions to harpacticoid systematics. As in previous Amendment Bulletins (Wells, 1978, 1979, 1981) the page numbers in parentheses are those of the original Keys (Wells, 1976).

Family Longipediidae

Longipedia spinulosa Itô, 1981 is added to the genus (see p. 12 and Wells, 1981).

Family Cerviniidae

1. Pontostratiotes minor and P. fontana, new species by Dinet (1981), P. pacificus and P. unisetosus, new species by Itô (1982b), and P. sixtorum mindanaoensis Itô, 1982b, all to genus codon in KGG 1 (p. 21).
2.Montagna (1981) redescribes the male of Cervinia magna, discovering previously unknown sexual dimorphism. Brotskaya (1963) made C. magna the type of her new monotypic genus, Pseudocervinia. Montagna believes that this is not justified and proposes that Pseudocervinia sink as a synonym of Cervinia. As a consequence of Montagna's description —
(a)amend the codon for Pseudocervinia in KGG 1 (p. 22) to read — d/d/u/3:3:3:3:/2:2:2:2 — Cervinia magna ♀,
(b)note that C. magna ♂ has the same codon in KGG 100 (p. 23) as C. bradyi.
3. Cervinia unisetosa Montagna, 1981 to codon for C. tenuiseta in KGG 100 (p. 23).
4. Expansicervinia glacieria Montagna, 1981 to the codon for Cervinia tenuicauda in KGG 100 (p. 23).
5. Pontostratiotes robustus Itô, 1982b requires a new codon in KGG 200 (p. 24) — 8/?/6:7:7:8/5:5:6:5/>abd.
6. Tonpostratiotes tenuipedalis Itô, 1982b requires a new codon in KGG 200 (p. 24) — 7/5/6:7:7:7:/5:5:6:3/>abd.
7.Montagna (1981) synonymizes Stratiopontotes with Ameliotes. Itô (1982b) disagrees but synonymizes Ameliotes with Herdmaniopsis.

Family Ectinosomatidae

1. Arenosetella panamensis and A macronychospina, new species by Mielke (1981b), to genus codon in KGG 1 (p. 28).
2. Halectinosoma perforatum Itô, 1981 to codon for H. canaliculatum and H. inhacae in KGG 100 (p. 29).
3. Halectinosoma otakoua and H. hydrofuge, new species by Wells, Hicks & Coull (1982), to genus codon in KGG 100 (p. 29).
4. Noodtiella tabogensis Mielke, 1981b to codon for N. hoodensis in KGG 500 (p. 35).

Family Harpacticidae

Harpacticus alevtinae Tschislenko, 1977 requires a new codon in KGG 100 (p. 45) — nor/9/1/?/4:5/0:5. Tschislenko does not state if there are modifications to the male P.2-P.4.

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Family Tisbidae

1. Tisbe japonica Ho, 1982 and T. coulli, T. ianthina, T. longipes and T. variana, all new species by Volkmann, 1979a, to genus codon in KGG 100 (p. 49).
2.Volkmann (1979c) gives a partial revision of Tisbe, with keys to the species dealt with. In this revision she —
(a)describes seven new species — T. trisetosa, elanitica, rampighera, perplexa, gigantea, denticulata and maraensis; all to genus codon in KGG 100 (p. 49).
(b)synonymizes T. wilsoni with T. gracilis and T. lancii with T. holothuriae,
(c)describes T. inflatiseta Volkmann, 1979 for the nomen nudum, Tisbe inflatiseta Fava & Volkman, 1975.
(d)removes T. ompacta, cluthae, elongata and racovitzai to species incertae sedis in the genus,
(e)resurrects Bathyidia for B. remota Farran. This genus has the same codon in KGG 100 (p. 49) as Tisbe.
As a consequence of this revision amend note b) to KGG 100 (p. 50) to read — "Care must be taken with these three genera. Bathyidia can be separated from most species of Tisbe by the form of the P.1 endopod (see Volkmann, 1979c). Scutellidium and Tisbe can be separated on the respective absence or presence of a seta on the A.2 basis".
3.Volkmann (1979b) revises Tisbella. As a consequence —
(a)delete Idyellopsis and Zosime from KGG 1 (p. 48),
(b)delete Tisbella timsae and T. pulchella from KGG 200 (p. 51),
(c)describes T. inflatiseta Volkmann, 1979 for the nomen nudum, Tisbe inflatiseta Fava & Volkmann, 1975.
(d)insert KGG as follows —
KGG 300 — characters
1.A.1♀
n = number of segments.
2.A.2 Exp.
n = number of segments.
3.P.2-P.4 Exp.3
n:n:n = number of setae and/or spines on P.2, P.3 and P.4 respectively.
4.P.2-P.4 Enp.3
n:n:n = number of setae and/or spines on P.2, P.3 and P.4 respectively.
5.P.2-P.4 Enp.2
n:n:n = number of setae on inner border of P.2, P.3 and P.4 respectively.
KGG 300
A.1♀
segs.		 A.2
Exp.
segs.		 P.2-P.4
Exp.3
Setae		 P.2-P.4
Enp.3
Setae		 P.2-P.4
Enp.2
Setae
8 4 7:8:8 5:6:5 2:1:1 Idyellopsis
7(-8?) 3 7:8:8 5:6:5 2:2:1 Tisbella pulchella
7 4 7:8:8 5:6:5 2:2:1 Tisbellaa)
7 3 7:8:8 4:5:4 1:1:1 Zosimeb)
7 3 7:8:8 4:4:4 1:1:1 Z. reyssi
7 3 7:7:7 4:4:4 1:1:1 Zosimec)
6 3 7:7:7 4:4:4 1:1:1 Zosimed)
6 3 6:6:6 4:3:3 1:1:1 Z. bathyalis
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a) Tisbella timsae, rosea Volkmann, 1979b, alba Volkmann, 1979b.
b) Zosime incrassata, bathybia.
c) Zosime valida, atlantica, paratypica.
d) Zosime typica, major, mediterranea, gisleni, bergensis, erythraea, paramajor.
4.As a consequence of Kunz's (1981) revision of the Paramesochridae the following must be deleted from Tisbidae KGG 1 (p.48) —
(a)codons for the two species of Tisbisoma,
(b)codon for Idyanthopsis psammophila,
(c)footnote c).

Family Porcellidiidae

Hicks (1982a) redescribes Porcellidium tristanense and describes three new species — P. algoense, P. laurencium and P. ulvum.

Family Peltidiidae

1.KGG 1 (p. 53): New codons are required for —
Altheutha roeae Hicks, 1982a — 2/3/1:1:1/1:l:l/d:d
Eupelte beckleyae Hicks, 1982a— 2/2/0:0:0/1:l:2/d:d
Altheuthellopsis corallina Humes, 1981b — 1/2/0:0:0/l:l:l/f:f.
2.KGG 100 (p. 55): Eupelte hexaseta Hicks, 1982a has 6 setae.
3. Altheutha langi a synonym of A. depressa according to Hicks (1982a).

Family Tegastidae

1. Tegastes acroporanus Humes, 1981a and T. cnidicus Humes, 1981b to genus codon in KGG 1 (p. 56).
2. Parategastes coetzeei Kunz, 1980 to genus codon in KGG 1 (p. 56).
3.Kunz (1980) follows Monard (1935) in raising Parategastes sphaericus var. similis Sewell, 1924 to full species status. This species is included in the genus codon in KGG 1 (p. 56).

Family Parastenheliidae

Parastenhelia megarostrum Wells, Hicks & Coull, 1982 is added to the genus (see p. 12); a key to the genus is given.

Family Diosaccidae

1. Amphiascoides golikovi Tschislenko, 1977 to codon for A. littoralis in KGG 1100 (P. 74).
2.Miscegenus heretaunga Wells, Hicks & Coull, 1982: Because the A.2 exopod can have two or three segments this new genus and species requires new codons as follows —
in KGG 110 (p. 74) — <Exp/se/7/5:5/2:5
in KGG 600 (p. 89) — 5:6:7/0:0:0/2:5/5:5/m.
3. Robertsonia curtisii Greenwood & Tucker, 1982 to codon for R. diademata and R. angolensis in KGG 200 (p. 77).
4. Stenhelia (D.) latioperculata Itô, 1981 requires a new codon in KGG 400 (p. 86) — 4/7:8:7/7/d:d/lss.
5.KGG 600 (p. 89): New codons are required for —
Amphiascoides koltuni Tschislenko, 1977 — 5?:6:7/0?:0:()/2:4/5:5/?
Diosaccus aff. dentatus Itô, 1982a — 7:8:8/0:0:0/1:4/6?/?
6. Schizopera elatensis Kahan & Bar-El, 1982 to genus codon in KGG 800 (p. 93).
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Family Ameiridae

1. Parapseudoleptomesochra italica Pesce & Petkovski. 1980 to codon for Parapseudoleptomesochrad) in KGG 1 (p. 102 as Nitocrella, but see Wells, 1978 p. 5).
2. Praeleptomesochra phreatica Pesce, 1981 requires a new codon in KGG 1 (p. 101) — 3:3/2:3:3/2:2:2/4/1 — and a note that the existing genus codon now applies only to P. africana, P. pygmaea and P. similis.
3. Ameira parascotti Tschislenko, 1977 requires a new codon in KGG 400 (p. 114) — 0:0:0/1:1:1/1:1:1/4:5:5/4:5/?
4. Nitocrella petkovskii and N. paceae, both new species by Pesce (1980), require new codons in KGG 700 (p. 120) —
4:4:5/1:1:1/1:2:1/3:4/? — N. petkovskii
4:4:6/1:1:1/1:1:1/3:4/? — N. paceae.

Family Paramesochridae

1.Becker & Kunz (1981) transfer Oniscopsis to family Tetragonicipitidae; as a consequence —
(a)delete codons to O. robinsoni (p. 124) and O. pauliani (p. 125) in KGG 1,
(b)character 5 in KGG 1 is now redundant.
2.Kunz (1981) describes several new taxa —
(a) Diarthrodella secunda pacifica to species codon in KGG 1 (p. 124),
(b) Paramesochra acutata hawaiiensis requires a new codon in KGG 100 (p. 126) — 4/4:4:2/0:0:0/1:3/0:3,
(c) Kliopsyllus insularis requires a new codon in KGG 200 (p. 127) — 4:4:2/1:1:1/2:3/1:4,
(d) Kliopsyllus debilis to codon for K. enalius in KGG 200 (p. 127),
(e) Kliopsyllus californicus to codon for Kliopsyllusc) in KGG 200 (p. 127),
(f) Kliopsyllus spiniger ornatus to species codon in KGG 200 (p. 127).
3.Kunz (1981) revises the family. As a consequence changes to KGG 1 (p. 124) are necessary to accommodate —
(a)the reduction of Paraleptopsyllus to a subgenus of Leptopsyllus, and Intermedop-syllus to a subgenus of Scottopsyllus,
(b)the placement of Kliopsyllus runtzi, K. abyssalis and K. gigas in Wellsopsyllus, a new subgenus of Scottopsyllus,
(c)Kunz's formal proposal that Idyanthopsis psammophila be transferred to Diarthrodella,
(d)the transfer of Tisbisoma to this family; add a new codon — 2-3:3:3:3/3/5/8/nf.
4. Paramesochra borealis Geddes, 1981a requires a new codon in KGG 100 (p. 126) — 4/4:4:2/0:1:1/2:3/2:4.
5.Amend the codons in KGG 100 (p. 126) for these species to read —
4/4:4:2/0:0:1/2:4/0:2 Paramesochra longicaudata
4/4:4:2/0:0:1/2:3/0:3 P. helgolandica
4/4:3:2/0:0:0/1:3/0:3 P. acutata s.str.
6.Amend the codon in KGG 300 (p. 129) for Apodopsyllus camptus to read — 1/5/0:4/0:4. On further examination it is clear to me that the "inner seta" of the P.5 basendopod is only a remnant of the inner lobe.

Family Tetragonicipitidae

1.Three new species of Phyllopodopsyllus by Kitazima (1981) need to be added to KGG 1 (p. 131) —
(a) P. simplex: female to codon for P. danielae; male to codon for P. bahamaensis and P. opisthoceratus♀.page 5
(b) P. punctatus: female to codon for P. mossmani and P. laspalmensis; male to codon for P. paramossmani♂.
(c) P. setouchensis: female to codon for P. bermudae et. al.; male requires a new codon — a/2vr/2/3:3:2/5:6:6.
2. Phyllopodopsyllus borutzkyi: Codon in KGG 1 (p. 132) refers to female only; add a new codon for the male — a/2wd/2/3:2:2/4:4:6.
3.Add a new codon to KGG 1 (p. 133) — a/a/2/2-3:1:1/3:2:1 — Oniscopsis.

Family Canthocamptidae

In KGG 100 (p. 138) amend the codon for Mesochra paranaensis to read — 6/3/7:7:7/5:5/5:5/?

Family Cylindropsyllidae

1. Boreopontia heipi Willems, 1981 to codon for Arenopontia australis in KGG 1 (p. 141).
2. Syrticola flandricus Willems & Claeys, 1982 requires a new codon in KGG 1 (p. 141) — s/2:na/f/1:1:1/p.
3.Geddes (1981a) redescribes the female of Leptastacus rostratus (the male remains unknown). As a consequence amend the species codon in KGG 300 (p. 144) to read — p/2/0:0:1/3:4:5/1:1:0/1:2:2.
4.Geddes (1981a) raises Leptastacus rostratus tourica to species status; it has the same codon in KGG 300 (p. 144) as L. rostratus.

Family Cletodidae

1.Thistle (1980) describes two new species of Enhydrosoma, reviews the genus, comments on several species and provides a key; as a consequence —
(a) Enhydrosoma franklini Thistle, 1980: Because segments 1-2 of P.2-P.4 exopod are fused together this species has the same codon in KGG 1 (p. 155) as Enhydrosomella, but the line of fusion is obvious and E. franklini cannot be confused with any species of Enhydrosomella,
(b) Enhydrosoma woodini Thistle, 1980 requires a new codon in KGG 500 (p. 165) — 2:3/a/d:3:2/d:2:2/♀,
(c)amendments to several codons in KGG 500 (p. 165) are required — 2:3/a/d:3:4/d:2-3:4/s — E. curticauda
2:2-3/a/d:3:5/d:2-3:5/m — E. curvirostre
2:2-3/a/d:3:4-5/d:2:2/♀ — E. propinquum
2:3/a/d:3:4/d:3:4/m-s — E. sarsi
2. Heteropsyllus pseudonunni Coull & Palmer, 1980 to codon for H. confluens in KGG 1 (p. 158).
3.New codons are required in KGG 1 (p. 154) for two new monotypic genera described by Apostolov (1980) —
3:2/3:3:3/2:2/5:5:6/3:3 — Pontocletodes ponticus
2:1/3:3:3/2:2/5:6:6/4:4 — Miroslauia longicaudata
4. Enhydrosoma variabile Wells, Hicks & Coull, 1982 requires a new codon in KGG 500 (p. 165) — 2:3/a/d:2:3-4/d:2:2/s.

Family Laophontidae

1. Heterolaophonte serratula Mielke, 1981a requires new codons —
in KGG 1-♀♀ (p. 172) — 3:3:2/2:2:1/5:6/2/7
in KGG 1-♂♂ (p. 188) — 3:3:2/2:l/5b)/2/2.
2. Stygolaophonte arenophila: Amend the codon in KGG 1-♀♀ (p. 173) to read —
3:3:2/2:2:1/4:2/2/7. page 6
3. Esola longicauda galapagoensis Mielke, 1981a: As the outer seta of ♀P.5 Benp. is very reduced and spinule-like it would be advisable to add a new codon in KGG 1-99 (p. 172) - 3:3:3/2:2:2/2(3?):6/2/6. Note also that the species codon in KGG 900-♀♀ refers only to the nominate subspecies. The male of this new subspecies keys out with the species codon in KGG 600-♂♂ (p. 195).
4. Afrolaophonte schmidti Mielke, 1981a requires new codons —
in KGG 1-♀♀ (p. 172) — 1:3:3/0:1:1/4:4/1/5
in KGG 1-♂♂ (p. 188) — l:3:3/0:1s/0:3/1/2.
5. Klieonychocamptoides arganoi Cottarelli & Mura, 1980 and K. itoi Mielke, 1981a both require the same new codons —
in KGG 1-♀♀ (p. 172) — 1:1:1/0:0:0/3:4/1/6
in KGG 1-♂♂ (p. 188) — l:l:l/0:0/7b)/l/l.
6. Klieonychocamptoides remanei: Amend codon in KGG 1-♀♀ (p. 172) to read —
1:1:1/0:0:0/3:4/1/7.
7. Mexicolaophonte arganoi Cottarelli, 1977 requires new codons —
in KGG 1-♀♀ (p. 172) — 2:2:2/0:2:2/5:5/1/6
in KGG 1-♂♂ (p. 188) — 2s:1s:2/0:l/0:4/l/2.
8.Geddes (1982) redescribes Laophontina dubia; amend present codons to read —
KGG 1-♀♀ (p. 172) — 2s:1:3/0:0:l/4:5/l/6
KGG 1-♂♂ (p. 188) — 2s:1:3/0:l/0:3/l/0.
9.Mielke (1981a) describes Galapalaophonte pacifica n.gen., n.sp. but later (1982) synonymizes the species with Laophontina triarticulata. As a consequence amend the codon in KGG 1-♀♀ (p. 172) to read —
1s-2s:3:3/2:2:1/4:5/1/6.
10. Paralaophonte aenigmaticum Wells, Hicks & Coull, 1982: Male to codon for Arenolaophonte stygia in KGG 1-♂♂ <5(p. 190). Variability in the P.5♀ causes this species to
(a)key out to Esola rosei in KGG 1-♀♀ (p. 173),
(b)require a new codon in KGG 500-♂♂ (p. 179)
— 6:5-6:4/1:1:1/0:0:0/4:5:3/0:0:0.
11. Quinquelaophonte Wells, Hicks & Coull, 1982 is erected for the quinquespinosa-group of Heterolaophonte. As a consequence name changes are required to Heterolaophonte in KGG 1400-♀♀ (p. 183) and KGG 1800-♂♂ (p. 185), and to H. parasigmoides, H. quinquespinosa, H. capillata, H. longifurcata and H. wellsi in footnotes h and i of KGG 1-♂♂ (p. 191).
12. Quinquelaophonte candelabrum Wells, Hicks & Coull, 1982 requires new codons in KGG 1-♀♀ (p. 172) — 3:3:3/2:2:2/5:5/2/5
in KGG 1-♂♂ (p. 188) — 3:3:3/2:2/4b)/2/2.
13.Wells, Hicks & Coull (1982) give keys to Paralaophonte and Quinquelaophonte.
14. Loureirophonte isabelensis Mielke, 1981a requires new codons —
in KGG 100-♀♀ (p. 177) — 5:5:4/1:l:0/0:0:0/3:3:2/0:0:na
in KGG 1-♂♂ (p. 188) — 3:3:3/2:1/1:5/2/2.
The present genus codon in KGG 1-♀♀ now applies only to L. catharinensis and L. paranaensis.
15. Echinolaophonte tetracheir Mielke: Female to codon for E. horrida and E. brevispinosa in KGG 300-♂♂ (p. 177). Male requires a new codon in KGG 200-♂♂ (p. 193) — 6:7:6/l:l:l/3:4:3/0:0/a.
16. Paralaophonte panamensis Mielke, 1982 requires new codons —
in KGG 500-♀♀ (p. 179) — 6:7:6/1:1:1/0:0:0/4:6:4/0:0:0
in KGG 1200-♂♂ (p. 197) — 6:5:5/1:1:l/4:4:4/0:0/p.
17. Paralaophonte innae Tschislenko, 1977 requires new codons —
in KGG 600-♀♀ (p. 179) — 6:7:7/0:0:0/4:5:4/0:0:0
in KGG 600-♂♂ p. 195) — 6:7:7/l:l:l/4:5:4/0:0/a. page 7
18. Paralaophonte pacifica galapagoensis Mielke, 1981a to species codons in KGG 700-♀♀ (p. 180) and KGG 1100-♂♂ (p. 197).
19. Paralaophonte problematica Mielke, 1981a: Male to codon for Paralaophontea) in KGG 1100-♂♂ (p. 197). Female requires a new codon in KGG 700-♀♀ (p. 180) — 6:7:7/1:1:1/0:0:0/4:5:4/0:0:0.
20. Laophonte galapagoensis Mielke, 1981a: Male to codon for Paronychocamptuse) in KGG 1-♂♂ (p. 189). Female requires a new codon in KGG 1200-♀♀(p. 182) — 6:7:7/1:1:1/0:0:0/4:6:4/0:0:1.
21.KGG 1600-♂♂ (p. 184): Delete the last codon in this KGG. The codon 6:7:7/1:1:1/0:1:1/7:6:6/1:1:1 now leads to Pseudocletopsyllus spiniger and four species of Cletopsyllus (bacescui, secundus, sakagamii and brattstroemi Geddes, 1981b). Male of C. brattstroemi to genus codon in KGG 1-♀♀ (p. 190).
22. Laophontina sp.♂ Mielke, 1982 requires a new codon in KGG 1-♂♂ (p. 188) —1:2:3/0:2/0:5/1/2.
23.Hicks (1982b) describes the true male of Laophonte danversae. As a consequence
(a)add a new codon to KGG 1-♂♂ (p. 188) — 3:3:3/2:2/2:5/3/2,
(b)delete the codon for this species in KGG 1500-♂♂ (p. 198).

Family Ancorabolidae

Paralaophontodes exopoditus Mielke, 1981a requires a new codon in KGG 1 (p. 202) — 2:2/0:0/4-5/5:5:5/na:na. Note that the existing genus codon now applies only to P. echinatus and P. robustus.

page 8

References

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Becker, K. H. & Kunz, H. , 1981. Die Stellung der Gattung Oniscopsis (Chappuis) im System der Harpacticoida (Crustacea, Copepoda). Mitteilungen aus dem Zoologischen Museum der Universitat Kiel 1 (8): 34-37.

Brotskaya. V. A. , 1963. A survey of the Family Cerviniidae (Crustacea, Copepoda). Zoologicheskii Zhurnal 42: 1785-1803 (in Russian).

Cottarelli V. , 1977. Mexicolaophonte arganoi n.gen. n.sp. di Laophontidae (Crustacea, Copepoda, Harpacticoida) di acque interstiziali litorali messicane. Accademia Nazionale dei Lincei. Problemi attuali di scienza e di cultura, sezione: Missioni ed Esplorazioni — 1 171: 91-99.

Cottarelli, V. & Mura, G. , 1980. Klieonychocamptoides arganoi n.sp., arpacticoide di acque interstiziali litorali della Isola Maldive (Crustacea, Copepoda). Cahiers de Biologie Marine 21: 355-361.

Coull, B. C. & Palmer, M. A. , 1980. Heteropsyllus (Copepoda, Harpacticoida): A revised key, including a new species from Chesapeake Bay. Transactions of the American Microscopical Society 99: 303-309.

Dinet, A. , 1981. Description de deux nouvelles especes abyssales de Pontostratiotes (Crustacea, Copepoda, Harpacticoida). Bulletin de la Societe Zoologique de France 106: 201-211.

Fava, G. & Volkmann, B. , 1975. Tisbe (Copepoda: Harpacticoida) species from the Lagoon of Venice. 1. Seasonal fluctuations and ecology. Marine Biology 30: 151-165.

Geddes. D. C. , 1981a. On two interstitial marine harpacticoids (Crustacea: Copepoda) from northern Norway. Sarsia 66: 19-24.

Geddes, D. C. , 1981b. Marine biological investigations in the Bahamas. 21. A new species of Cletopsyllus (Copepoda, Harpacticoida). Sarsia 66: 287-291.

Geddes, D. C. , 1982. A redescription of Laophontina dubia Norman & T. Scott (Crustacea: Copepoda: Harpacticoida). Zoological Journal of the Linnean Society of London 74: 105-109.

Greenwood, J. C. & Tucker, M. J. , 1982. A new species of Robertsonia (Copepoda, Harpacticoida) from Port Curtis, Queensland. Crustaceana 42: 288-294.

Hicks, G. R. F. , 1982a. Porcellidiidae and Peltidiidae (Copepoda: Harpacticoida) from the marine algae of St Croix Island, Algoa Bay, South Africa. Zoological Journal of the Linnean Society of London 75: 49-90.

Hicks, G. R. F. , 1982b. New records of harpacticoid copepods from the east coast of Britain, and a description of the true male of Laophonte danversae Hamond. Crustaceana 42: 302-307.

Ho, J. S. , 1982. Copepoda associated with echinoderms of Japan. Annual Report of the Sado Marine Biological Station, Niigata University 12: 33-61.

Humes, A. G. , 1981a. A new species of Tegastes (Copepoda: Harpacticoida) associated with a scleractinian coral at Enewetok Atoll. Proceedings of the Biological Society of Washington 94: 254-263.

Humes, A. G. , 1981b. Harpacticoid copepods associated with Cnidaria in the Indo-West Pacific. Journal of Crustacean Biology 1: 227-240.

Itô, T. , 1981. Descriptions and records of marine harpacticoid copepods from Hokkaido, VIII. Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology 22: 422-450.

Itô, T. , 1982a. Diosaccus sp. aff. dentatus (Thompson et A. Scott) (Copepoda, Harpacticoida) from Mactan [sl., The Philippines. Publications of the Seto Marine Biological Laboratory 27: 165-171.

Itô, T. , 1982b. Harpacticoid copepods from the Pacific abyssal off Mindanao. I. Cerviniidae. Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology 23: 63-127.

Kahan, D. & Bar-El, T. , 1982. Schizopera elatensis n.sp. (Copepoda, Harpacticoida) from Elat, Israel. Crustaceana 42: 308-314.

Kitazima, Y. , 1981. Three new species of the genus Phyllopodopsyllus (Copepoda, Harpacticoida) from the Inland Sea of Japan. Publications of the Seto Marine Biological Laboratory 26: 393-424.

Kunz, H. , 1980. A new species of Parategastes (Copepoda, Harpacticoida) from South Africa. South African Journal of Zoology 15: 186-189.

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Kunz, H. , 1981. Beitrag zur Systematik der Paramesochridae (Copepoda, Harpacticoida) mit Beschreibung einiger neuer Arten. Mitteilungen aus dem Zoologischen Museum der Universität Kiel 1 (8): 2-33.

Mielke, W. , 1981a. Interstitielle Fauna von Galapagos. XXVIII. Laophontinae (Laophontidae), Ancorabolidae (Harpacticoida). Akademie der Wissenschaften und der Literatur, Mainz, mathematisch-naturwissenschaftlichen Klasse. Mikrofauna des Meeresbodens 84: 1-104.

Mielke, W. , 1981b. Interstitielle Ectinosomatidae (Copepoda) von Panama. Akademie der Wissenschaften und der Literatur, Mainz, mathematisch-naturwissenschaftlichen Klasse. Mikrofauna des Meeresbodens 85: 1-45.

Mielke, W. , 1982. Einige Laophontidae (Copepoda. Harpacticoida) von Panama. Crustaceana 42: 1-11.

Monard, A. , 1935. Les harpacticoides marins de la region de Salammbo. Bulletin Station Oceanographique de Salammbo 34: 3-94.

Montagna, P. A. , 1981. A new species and a new genus of Cerviniidae (Copepoda: Harpacticoida) from the Beaufort Sea, with a revision of the family. Proceedings of the Biological Society of Washington 93: 1204-1219.

Pesce, G. L. , 1980. Two new species of phreatic harpacticoids from Iran (Crustacea: Copepoda). Bijdragen tot de Dierkunde 50: 364-368.

Pesce, G. L. , 1981. A new harpacticoid from phreatic waters of Morocco, and remarks on the genus Praeleptomesochra Lang (Crustacea: Copepoda: Ameiridae). Bulletin Zoologisch Museum Universiteit van Amsterdam 8: 69-72.

Pesce, G. L. & Petkovski, T. K. , 1980. Parapseudoleptomesochra italica n.sp., a new harpacticoid from subterranean waters of Italy (Crustacea, Copepoda, Ameiridae). Fragmenta Balcanica 11: 33-42.

Thistle, D. , 1980. Enhydrosoma (Copepoda, Harpacticoida): An update and two new species. Transactions of the American Microscopical Society 99: 384-397.

Tschislenko, L. L. , 1977. Harpacticids (Copepoda, Harpacticoidea) from sponges of Franz Josef Land. Issledovaniia fauny morei: Akademiia nauk SSSR, Zoologischeskii Institut, Moscow 14 (22): 237-276 (in Russian).

Volkmann, B. , 1979a. Tisbe (Copepoda, Harpacticoida) species from Bermuda and zoogeographical considerations. Archivio di Oceanografia e Limnologia 19 supplement: 1-76.

Volkmann, B. , 1979b. A revision of the genus Tisbella (Copepoda, Harpacticoida). Archivio di Oceanografia e Limnologia 19 supplement: 77-119.

Volkmann, B. , 1979c. A revision of the genus Tisbe (Copepoda, Harpacticoida). Part 1. Archivio di Oceanografia e Limnologia 19 supplement: 121-284.

Wells, J. B. J. , 1976. Keys to Aid in the Identification of Marine Harpacticoid Copepods. Department of Zoology, University of Aberdeen , pp.215.

Wells, J. B. J. , 1978. Keys to Aid in the Identification of Marine Harpacticoid Copepods. Amendment Bulletin No. 1. Zoology Publications from Victoria University of Wellington 70:1-11.

Wells, J. B. J. , 1979. Keys to Aid in the Identification of Marine Harpacticoid Copepods. Amendment Bulletin No. 2. Zoology Publications from Victoria University of Wellington 73: 1-8.

Wells, J. B. J. , 1981. Keys to Aid in the Identification of Marine Harpacticoid Copepods. Amendment Bulletin No. 3. Zoology Publications from Victoria University of Wellington 75: 1-13.

Wells, J. B. J. , Hicks, G. R. F. & Coull, B. C. , 1982. Common harpacticoid copepods from New Zealand harbours and estauries. New Zealand Journal of Zoology 9: 151-184.

Willems, K. A. , 1981. Boreopontia heipi n.g., n.sp. an interstitial harpacticoid (Copepoda) from the southern bight of the North Sea. Biologisch Jaarboek Dodonaea, 49: 200-209.

Willems, K. A. & Claeys, D. , 1982. Syrticola flandricus n.g., n.sp., a harpacticoid copepod from the southern bight of the North Sea. Crustaceana 43: 1-8.

Professor J. B. J. Wells,
Department of Zoology,
Victoria University of Wellington,
Private Bag,
Wellington,
New Zealand.

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